Yun Yayao secara tidak sengaja melakukan perjalanan ke sebuah novel di mana ia menjadi karakter yang dapat dihabiskan yang meninggal pada awalnya. Setelah membunuh tiran, Tu Shan Jin, jiwa mereka bertukar. Ketika mereka menavigasi serangkaian krisis bersama, mereka mengembangkan perasaan satu sama lain dan akhirnya berakhir bersama.

Tanjiro Kamado, joined with Inosuke Hashibira, a boy raised by boars who wears a boar's head, and Zenitsu Agatsuma, a scared boy who reveals his true power when he sleeps, board the Infinity Train on a new mission with the Flame Pillar, Kyojuro Rengoku, to defeat a demon who has been tormenting the people and killing the demon slayers who oppose it!

Tanjiro Kamado, joined with Inosuke Hashibira, a boy raised by boars who wears a boar's head, and Zenitsu Agatsuma, a scared boy who reveals his true power when he sleeps, board the Infinity Train on a new mission with the Flame Pillar, Kyojuro Rengoku, to defeat a demon who has been tormenting the people and killing the demon slayers who oppose it!